[WING] WING: Pale or dark spots - Presence [WING] WING: 2nd rad cell, covered by pale spot, costal-tip part [WING] WING: r3 and m1, pale spots, distal part - Presence [WING] WING: r3 and m1, pale spots, distal part - Size [WING] [HEAD] WING: r3 and m1, pale spots, distal part - Connection [WING] WING: r3 and m1, pale spots distal part - Position [WING] WING: m2, pale spot, distal part - Presence [WING] WING: m1, pale spot, from proximal to median part - Presence [WING] [HEAD] WING: m1, pale spot layer and cross the veins M1 and M2 - Presence [WING] WING: m2, pale spot/area, from proximal to median part - Presence [WING] WING: m2, pale spot over r-m crossvein fused with the m2 spot which layers and crosses vein M2 - Presence [WING] WING: m, pale spot/area - Presence [WING] WING: r3, 4th pale costal spot versus 3rd dark costal spot - Size [WING] WING: r3, area of 4th p.c.s. bigger than 3rd d.c.s. - Shape of the 3rd d.c.s. [WING] WING: anal cell, pale spot in distal part - Presence [WING] WING: cua1, center spot - Presence and Colour [WING] WING: r-m crossvein, dark spot in the corner with M1 vein- Presence [WING] WING: arculus, dark spot under arculus - Presence [WING] WING: M1, pale spot/band spanning the vein - Presence [WING] WING: M1, pale spot in the median part - Position [WING] WING: M2, pale spot/band spanning the vein - Presence [WING] WING: M1, M2 and CuA1, at least 1 pale spot/area, abuts wing margin, apex of veins in distal part - Presence [WING] WING: M1, M2 and CuA1, pale spots surrounded by dark area, apex of veins - Shape [WING] WING: M2,dark spot in proximal part - Shape [WING] WING: Pale wing with only 2 dark areas on CuA1 and 2nd rad cell - Presence [WING] WING: m and anal cells, macrotrichia abundance - Presence [ABDOMEN] ABDOMEN: Spermathecae - Number [ABDOMEN] ABDOMEN: Spermathecae, sclerotized ring at the end of the spermathecal duct - Presence [ABDOMEN] ABDOMEN: Spermathecae, sclerotized ring at the end of the spermathecal duct - Shape [ABDOMEN] ABDOMEN: 1 or 2 spermathecae, pigmented neck - Presence [ABDOMEN] ABDOMEN: 1 Spermatheca - Shape [ABDOMEN] ABDOMEN: 1 spermatheca, curved shape - Presence [ABDOMEN] ABDOMEN: 1 spermatheca, spermathecal duct swollen - Presence [ABDOMEN] ABDOMEN: 1 spermatheca, spermathecal duct - Length [ABDOMEN] ABDOMEN: 2 spermathecae - Shape [ABDOMEN] ABDOMEN: Spermathecae, abdominal sclerites - Presence [ABDOMEN] ABDOMEN: Spermathecae, abdominal sclerites - Shape [ABDOMEN] ABDOMEN: 2 spermathecae - Size [ABDOMEN] ABDOMEN: 3 spermathecae - Shape [ABDOMEN] ABDOMEN: 3 spermathecae - Texture [LEG] LEG: forelegs, spines on tarsal segments - Presence [LEG] LEG: middle legs, spines on tarsal segments - Presence [LEG] LEG: hind legs, spines on tarsal segments - Presence [HEAD] EYES: interfacetal hairs - Presence [HEAD] EYES: Inter-ocular space - Shape [HEAD] MANDIBLE/MAXILLE: teeth - Presence [HEAD] CIBARIAL ARMATURE: cibarial armature - Presence [HEAD] PHARYNX POSTERIOR ARMATURE: pharynx posterior armature - Presence [HEAD] PALP: 3rd palpal segment - Shape [HEAD] PALP: 3rd palpal segment, sensory pits - Number [HEAD] PALP: 3rd palpal segment, single sensory pit - Opening versus depth [HEAD] ANTENNA: short segments - Shape [HEAD] ANTENNA: sensilla coeloconica, short segments - Presence [HEAD] ANTENNA: short sensilla trichodea, distal part segments IV to X - Number [HEAD] ANTENNA: long sensilla trichodea, proximal segments III-X - Shape [HEAD] ANTENNA: antennal XI/X ratio, length of segment XI divided by length of segment X - Range [HEAD] ANTENNA: sensilla coeloconica, segments III to VI - Presence [HEAD] ANTENNA: sensilla coeloconica, segments VII to X - Presence ANTENNA: sensilla coeloconica, segments XI to XV - Presence [WING] WING: anal cell, dark area abuts wing margin - Presence
achrayi 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined not applicablenot applicablenot applicableabsence no spot, homogenous cell not applicableno spot, homogenous cell not applicableno spot, homogenous cell absence of at least one of two not applicableabsence uniform absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel short ; long not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen multiple irregular pits not applicableflask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV not applicable
alazanicus 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined not applicablenot applicablenot applicablepresence no spot, homogenous cell not applicableno spot, homogenous cell not applicablefused with the r-m crossvein spot or 1 spot absence of at least one of two ; area of pale c.s. smaller than dark c.s. or approximatively equals not applicable1 pale spot clearly separated or abuted with wing margin 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel absence not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit inflated ?2 setae each classical > 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X presence on segments XI to XV not applicable
albicans No pale spot cover the r-m crossvein not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 3rd tarsomere no spines absence separated narrowly (1 or 2 facets) ; joined for a short distance (1 or 2 facets) absence both on madibles and maxilles absence absence triangular and moderately swollen multiple irregular pits not applicableflask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV not applicable
albihalteratus 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined not applicablenot applicablenot applicableabsence no spot, homogenous cell not applicableno spot, homogenous cell not applicableno spot, homogenous cell absence of at least one of two not applicableabsence uniform absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 3rd tarsomere no spines absence separated narrowly (1 or 2 facets) absence on maxilles and presence on mandibles absence absence triangular and moderately swollen single narrow opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XIII to XV not applicable
algeriensis 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined not applicablenot applicablenot applicableabsence no spot, homogenous cell not applicableno spot, homogenous cell not applicableno spot, homogenous cell absence of at least one of two not applicableabsence uniform absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel absence ; short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X presence on segments XI to XV not applicable
azerbajdzhanicus No pale spot cover the r-m crossvein not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence conical long not applicablenot applicablenot applicablenot applicableoval absence not applicableunequal not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence presence triangular and moderately swollen single wide opening and shallow pit inflated ?1 seta each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X absence on segments XI to XV not applicable
begueti 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined not applicablenot applicablenot applicableabsence no spot, homogenous cell not applicableno spot, homogenous cell not applicableno spot, homogenous cell absence of at least one of two not applicableabsence uniform absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 3rd tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence strongly swollen single narrow opening and deep pit flask shape ?1 seta each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X presence on segments XI to XV not applicable
begueti variation 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined not applicablenot applicablenot applicableabsence no spot, homogenous cell not applicableno spot, homogenous cell not applicableno spot, homogenous cell absence of at least one of two not applicableabsence uniform absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 3rd tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence strongly swollen single narrow opening and deep pit flask shape ?1 seta each classical < = 2 presence on segments III, IV, V and VI presence on segments VII and IX presence on segments XI to XV not applicable
brunnicans No pale spot cover the r-m crossvein ; 1 or more pale spots less than 1/3 pale ; more than 1/3 No pale spots or areas badly defined not applicablenot applicablenot applicableabsence no spot, homogenous cell not applicableno spot, homogenous cell not applicableno spot, homogenous cell absence of at least one of two not applicableabsence uniform absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicablepresence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen multiple irregular pits not applicableflask shape ?2 setae each classical < = 2 presence on segments III and V presence on segments VII and IX presence on segments XI to XV not applicable
brunnicans variation No pale spot cover the r-m crossvein ; 1 or more pale spots less than 1/3 pale ; more than 1/3 No pale spots or areas badly defined not applicablenot applicablenot applicableabsence no spot, homogenous cell not applicableno spot, homogenous cell not applicableno spot, homogenous cell absence of at least one of two not applicableabsence uniform absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicablepresence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen multiple irregular pits not applicableflask shape ?2 setae each classical < = 2 presence on segments III, V and VI ; presence on segments III, IV, V and VI presence on segments VII, VIII and IX ; presence on segments VIII, IX and X ; presence on segments VII, VIII, IX and X presence on segments XI to XV not applicable
cameroni No pale spot cover the r-m crossvein not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableabsence presence 2 functional and 1 rudimentary absence not applicableshort not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicablespines on 1st and 2nd tarsomere ; spines on 1st, 2nd and 3rd tarsomere spines on 1st to 3rd tarsomere ; spines on 1st to 4th tarsomere spines on 1st and 2nd tarsomere ; spines on 1st to 3rd tarsomere absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single narrow opening and shallow pit inflated ?1 seta each classical < = 2 ; > 2 presence on segment III absence on segments VII to X absence on segments XI to XV not applicable
cataneii or gejgelensis variation 1 or more pale spots wholly dark, homogenous Pale spots well defined sizes approximatively equal separated broadly abut wing margin presence 1 spot well defined absence pale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. smaller than dark c.s. or approximatively equals not applicable1 pale spot clearly separated or abuted with wing margin 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII and IX presence on segments XI to XV not applicable
caucoliberensis 1 or more pale spots wholly dark, homogenous ; less than 1/3 pale Pale spots well defined sizes approximatively equal fused broadly abut wing margin presence 1 spot well defined absence pale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. smaller than dark c.s. or approximatively equals not applicable1 pale spot clearly separated or abuted with wing margin 1 pale spot surrounded by dark area absence absence 1 pale spot in the median part of M1 m1 spot more distal than M1 spot ; M1 and costal spots fused absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary ??short not applicablenot applicablenot applicablenot applicableoval absence not applicableunequal not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen multiple irregular pits not applicableflask shape ?2 setae each classical < = 2 presence on segments III and VI presence on segments VII, VIII, IX and X presence on segments XI to XIV not applicable
chiopterus 1 or more pale spots more than 1/3 No pale spots or areas badly defined not applicablenot applicablenot applicableabsence ; presence no spot, homogenous cell ; undefined spot/area not applicableno spot, homogenous cell ; pale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot absence of at least one of two ; area of pale c.s. smaller than dark c.s. or approximatively equals not applicable1 pale spot clearly separated or abuted with wing margin uniform ; 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence absence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals ; unequal not applicablenot applicableno spines spines on 1st to 3rd tarsomere no spines scattered and short hairs joined for a short distance (1 or 2 facets) presence both madibles and maxilles absence absence slender or slightly swollen ; triangular and moderately swollen single narrow opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV not applicable
circumscriptus 1 or more pale spots wholly dark, homogenous Pale spots well defined r3 spot clearly larger than m1 spot separated separated from wing margin presence 1 spot well defined presence 1 spot ; pale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. bigger than dark c.s. anterior, median and posterior margin approximatively equals 2 pale spots fused ; 2 pale spots separated 1 pale spot surrounded by dark area presence absence almost a wide range of M1 pale not applicablealmost a wide range of M2 pale presence presence not describedabsence presence 1 functional absence not applicableshort oval absence absence ?not applicableabsence not applicablenot applicablenot applicablenot applicableno spines spines on 1st to 3rd tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence presence triangular and moderately swollen single wide opening and deep pit flask shape ?2 setae each classical < = 2 ; > 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X presence on segments XI to XIV ?
clastrieri 1 or more pale spots wholly dark, homogenous Pale spots well defined sizes approximatively equal ; r3 spot clearly larger than m1 spot separated broadly abut wing margin presence 1 spot well defined absence pale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. smaller than dark c.s. or approximatively equals not applicable2 pale spots separated 1 pale spot surrounded by dark area absence absence 1 pale spot in the median part of M1 m1 spot more proximal than M1 spot ; M1 and costal spots separated absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableunequal not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence presence triangular and moderately swollen single wide opening and shallow pit flask shape ?2 setae each classical < = 2 ; > 2 presence on segment III absence on segments VII to X presence on segments XI to XV not applicable
clintoni 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined ; Pale spots well defined sizes approximatively equal separated broadly abut wing margin presence undefined spot/area not applicablepale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. smaller than dark c.s. or approximatively equals not applicable1 pale spot clearly separated or abuted with wing margin ; 2 pale spots fused 1 pale spot surrounded by dark area absence absence 1 pale spot in the median part of M1 M1 and costal spots fused ; M1 and costal spots separated 1 pale spot in median part of M2 absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableunequal not applicablenot applicableno spines spines on 1st to 3rd tarsomere no spines absence separated narrowly (1 or 2 facets) absence on maxilles and presence on mandibles absence absence slender or slightly swollen ; triangular and moderately swollen single narrow opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XIII to XV not applicable
comosioculatus No pale spot cover the r-m crossvein not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel absence not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines dense and long hairs separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and deep pit flask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV not applicable
corsicus No pale spot cover the r-m crossvein not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence donut long not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals ; unequal not applicablenot applicableno spines spines on 1st to 3rd tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit inflated ?1 seta each classical < = 2 ; > 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X absence on segments XI to XV not applicable
deltus 1 or more pale spots more than 1/3 Pale spots well defined r3 spot clearly larger than m1 spot separated broadly abut wing margin presence undefined spot/area ; 1 spot well defined absence no spot, homogenous cell ; 1 spot not applicablefused with the r-m crossvein spot or 1 spot area of pale c.s. bigger than dark c.s. anterior, median and posterior margin approximatively equals 2 pale spots fused 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot describedabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence joined for a long distance (more than 2 facets) presence both madibles and maxilles absence absence slender or slightly swollen multiple irregular pits not applicableflask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV ?
derisor No pale spot cover the r-m crossvein not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel absence not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single narrow opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV not applicable
desertorum 1 or more pale spots less than 1/3 pale Pale spots well defined r3 spot clearly larger than m1 spot ?separated from wing margin presence 1 spot well defined ?pale spot over r-m crossvein fused with the m2 spot ?fused with the r-m crossvein spot or 1 spot area of pale c.s. bigger than dark c.s. median margin more narrow than anterior or posterior margin 1 pale spot clearly separated or abuted with wing margin 1 pale spot surrounded by dark area absence absence absence ?absence, no pale spot absence not applicablenot describedabsence presence 1 functional absence not applicableabsence ; short oval absence absence ?not applicableabsence not applicablenot applicablenot applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence presence triangular and moderately swollen single wide opening and shallow pit flask shape absence on at least one short segment 1 seta each classical < = 2 presence on segments III and V ; presence on segments III and VI ; presence on segments III, V and VI ; presence on segments III, IV, V and VI presence on segments VII and IX ; presence on segments VII, VIII and IX presence on segments XI to XIV ?
dewulfi 1 or more pale spots more than 1/3 No pale spots or areas badly defined not applicablenot applicablenot applicablepresence undefined spot/area ; 1 spot well defined absence pale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot absence of at least one of two ; area of pale c.s. smaller than dark c.s. or approximatively equals not applicable1 pale spot clearly separated or abuted with wing margin 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence absence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableunequal not applicablenot applicableno spines spines on 1st to 3rd tarsomere no spines absence joined for a short distance (1 or 2 facets) presence both madibles and maxilles absence absence slender or slightly swollen ; triangular and moderately swollen single narrow opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV not applicable
duddingstoni 1 or more pale spots wholly dark, homogenous Pale spots well defined sizes approximatively equal separated broadly abut wing margin presence 1 spot well defined absence pale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. smaller than dark c.s. or approximatively equals not applicable1 pale spot clearly separated or abuted with wing margin ; 2 pale spots fused 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel absence not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and deep pit flask shape ?2 setae each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X presence on segments XI to XV not applicable
dzhafarovi No pale spot cover the r-m crossvein not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel absence not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 3rd tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segment III presence on segments VII and IX presence on segments XI to XIV not applicable
faghihi No pale spot cover the r-m crossvein not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableabsence presence 2 functional and 1 rudimentary absence not applicableshort not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals ; unequal not applicablenot applicablespines on 1st and 2nd tarsomere spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape presence on all short segments 1 seta each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X absence on segments XI to XV not applicable
fagineus 1 or more pale spots more than 1/3 Pale spots well defined r3 spot clearly larger than m1 spot separated broadly abut wing margin presence 1 spot well defined absence pale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. bigger than dark c.s. anterior, median and posterior margin approximatively equals 1 pale spot clearly separated or abuted with wing margin 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot describedabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence joined for a long distance (more than 2 facets) presence both madibles and maxilles presence absence slender or slightly swollen multiple irregular pits not applicableflask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV ?
fascipennis 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined not applicablenot applicablenot applicableabsence no spot, homogenous cell not applicableno spot, homogenous cell not applicablefused with the r-m crossvein spot or 1 spot absence of at least one of two not applicableabsence ; 1 pale spot clearly separated or abuted with wing margin uniform ; 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen multiple irregular pits not applicableflask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV not applicable
festivipennis 1 or more pale spots wholly dark, homogenous Pale spots well defined sizes approximatively equal ; r3 spot clearly larger than m1 spot separated broadly abut wing margin presence 1 spot well defined absence 1 spot not applicable2 or 3 spots separated area of pale c.s. smaller than dark c.s. or approximatively equals not applicable2 pale spots separated 1 pale spot surrounded by dark area absence absence 1 pale spot in the median part of M1 m1 spot more proximal than M1 spot ; M1 and costal spots separated absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableunequal not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence presence triangular and moderately swollen single wide opening and shallow pit flask shape ?2 setae each classical < = 2 ; > 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X presence on segments XI to XV not applicable
flavipulicaris 1 or more pale spots more than 1/3 Pale spots well defined r3 spot clearly larger than m1 spot fused broadly abut wing margin presence 1 spot well defined absence 1 spot not applicablefused with the r-m crossvein spot or 1 spot area of pale c.s. bigger than dark c.s. anterior, median and posterior margin approximatively equals 1 pale spot clearly separated or abuted with wing margin 1 dark spot surrounded by pale area absence absence absence absence, no pale spot absence, no pale spot presence absence large shape, elongated spot absence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence joined for a long distance (more than 2 facets) presence both madibles and maxilles absence absence slender or slightly swollen multiple irregular pits not applicableflask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV ?
furcillatus 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined not applicablenot applicablenot applicableabsence no spot, homogenous cell not applicableno spot, homogenous cell not applicableno spot, homogenous cell absence of at least one of two not applicableabsence uniform absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary absence not applicableshort not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single narrow opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segment III presence on segments VII and IX ; presence on segments VII, VIII and IX ; presence on segments VIII, IX and X ; presence on segments VII, VIII, IX and X presence on segments XI to XV not applicable
furcillatus variation 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined not applicablenot applicablenot applicableabsence no spot, homogenous cell not applicableno spot, homogenous cell not applicableno spot, homogenous cell absence of at least one of two not applicableabsence uniform absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary absence not applicableshort not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single narrow opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV not applicable
gejgelensis or cataneii variation 1 or more pale spots wholly dark, homogenous Pale spots well defined sizes approximatively equal separated broadly abut wing margin presence 1 spot well defined absence pale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. smaller than dark c.s. or approximatively equals not applicable1 pale spot clearly separated or abuted with wing margin 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X presence on segments XI to XV not applicable
griseidorsum 1 or more pale spots less than 1/3 pale ; more than 1/3 Pale spots well defined sizes approximatively equal separated broadly abut wing margin presence 1 spot well defined absence 1 spot ; pale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. smaller than dark c.s. or approximatively equals not applicable1 pale spot clearly separated or abuted with wing margin ; 2 pale spots fused 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel absence not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals ; unequal not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen multiple irregular pits not applicableflask shape ?2 setae each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X presence on segments XI to XIV not applicable
grisescens 1 or more pale spots more than 1/3 No pale spots or areas badly defined ; Pale spots well defined r3 spot clearly larger than m1 spot separated broadly abut wing margin presence undefined spot/area not applicablepale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. bigger than dark c.s. anterior, median and posterior margin approximatively equals 1 pale spot clearly separated or abuted with wing margin 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot describedabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicablespines on 1st and 2nd tarsomere spines on 1st to 4th tarsomere spines on 1st and 2nd tarsomere absence joined for a short distance (1 or 2 facets) presence both madibles and maxilles absence absence slender or slightly swollen multiple irregular pits not applicableflask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV ?
haranti 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined not applicablenot applicablenot applicablepresence no spot, homogenous cell not applicableno spot, homogenous cell not applicableno spot, homogenous cell absence of at least one of two ; area of pale c.s. smaller than dark c.s. or approximatively equals not applicable1 pale spot clearly separated or abuted with wing margin 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel absence ; short not applicablenot applicablenot applicablenot applicableoval absence not applicableunequal not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence strongly swollen single narrow opening and deep pit flask shape ?2 setae each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X presence on segments XI to XV not applicable
haranti variation 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined not applicablenot applicablenot applicablepresence no spot, homogenous cell not applicableno spot, homogenous cell not applicableno spot, homogenous cell absence of at least one of two ; area of pale c.s. smaller than dark c.s. or approximatively equals not applicable1 pale spot clearly separated or abuted with wing margin 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel absence ; short not applicablenot applicablenot applicablenot applicableoval absence not applicableunequal not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence strongly swollen single narrow opening and deep pit flask shape ?2 setae each classical < = 2 presence on segments III, IV, V and VI presence on segments VII and IX presence on segments XI to XIV ; presence on segments XI to XV not applicable
heliophilus No pale spot cover the r-m crossvein not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles presence absence triangular and moderately swollen single wide opening and shallow pit inflated ; flask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV not applicable
helveticus Pale wing, only 1 dark spot not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicablenot applicablepresence 1 functional presence parallel absence oval ; elongated finger shape appendage at the bottom absence ?not applicableabsence not applicablenot applicablenot applicablenot applicablespines on 1st, 2nd and 3rd tarsomere spines on 1st to 3rd tarsomere spines on 1st to 3rd tarsomere absence separated widely (more than 2 facets) ; separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence slender or slightly swollen multiple irregular pits not applicableflask shape ?1 seta each classical < = 2 presence on segments III, V and VI presence on segments VII, VIII, IX and X absence on segments XI to XV not applicable
heteroclitus 1 or more pale spots wholly dark, homogenous ; less than 1/3 pale No pale spots or areas badly defined not applicablenot applicablenot applicableabsence no spot, homogenous cell not applicableno spot, homogenous cell not applicableno spot, homogenous cell absence of at least one of two not applicableabsence ; 1 pale spot clearly separated or abuted with wing margin uniform ; 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary absence not applicableshort not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X presence on segments XI to XV not applicable
ibericus No pale spot cover the r-m crossvein not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableabsence presence 3 functional absence not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableAll spermathecae with hook-shaped apical processes - All processes long and with same length scale surface spines on 1st to 4th tarsomere spines on 1st to 4th tarsomere ?absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segment III presence on segments VII, VIII, IX and X absence on segments XI to XV not applicable
imicola 1 or more pale spots more than 1/3 Pale spots well defined r3 spot clearly larger than m1 spot separated broadly abut wing margin presence 1 spot well defined absence pale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. bigger than dark c.s. median margin more narrow than anterior or posterior margin 2 pale spots fused 1 pale spot surrounded by dark area absence absence almost a wide range of M1 pale not applicableabsence, no pale spot ; 1 pale spot in median part of M2 absence not applicablenot describedabsence absence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableunequal not applicablenot applicableno spines spines on 1st to 3rd tarsomere no spines absence joined for a short distance (1 or 2 facets) presence both madibles and maxilles absence absence slender or slightly swollen ; triangular and moderately swollen single ; multiple irregular pits narrow opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV ?
impunctatus 1 or more pale spots more than 1/3 No pale spots or areas badly defined ; Pale spots well defined r3 spot clearly larger than m1 spot separated broadly abut wing margin presence undefined spot/area not applicablepale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. bigger than dark c.s. anterior, median and posterior margin approximatively equals 1 pale spot clearly separated or abuted with wing margin 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot describedabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals ; unequal not applicablenot applicableno spines spines on 1st to 3rd tarsomere no spines absence joined for a short distance (1 or 2 facets) ; joined for a long distance (more than 2 facets) presence both madibles and maxilles absence absence slender or slightly swollen multiple irregular pits not applicableflask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV ?
indistinctus 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined not applicablenot applicablenot applicableabsence no spot, homogenous cell not applicableno spot, homogenous cell not applicableno spot, homogenous cell absence of at least one of two not applicableabsence uniform absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary absence not applicableshort not applicablenot applicablenot applicablenot applicableoval absence not applicableunequal not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence strongly swollen single narrow opening and deep pit flask shape ?2 setae each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X presence on segments XI to XIV not applicable
jumineri 1 or more pale spots wholly dark, homogenous ; less than 1/3 pale No pale spots or areas badly defined not applicablenot applicablenot applicablepresence undefined spot/area not applicable1 spot ; pale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. smaller than dark c.s. or approximatively equals not applicable1 pale spot clearly separated or abuted with wing margin 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel absence ; short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals ; unequal not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segments III and V presence on segments VII, VIII and IX presence on segments XI to XV not applicable
jumineri near bahrainensis 1 or more pale spots wholly dark, homogenous ; less than 1/3 pale No pale spots or areas badly defined not applicablenot applicablenot applicableabsence undefined spot/area not applicablepale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot absence of at least one of two not applicable1 pale spot clearly separated or abuted with wing margin 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel absence ; short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals ; unequal not applicablenot applicableno spines ?no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segments III and V presence on segments VII and IX presence on segments XI to XV not applicable
jurensis No pale spot cover the r-m crossvein not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel absence not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence strongly swollen single wide opening and deep pit flask shape ?2 setae each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X presence on segments XI to XV not applicable
kibunensis 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined not applicablenot applicablenot applicableabsence no spot, homogenous cell not applicableno spot, homogenous cell not applicableno spot, homogenous cell absence of at least one of two not applicableabsence uniform ; 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit inflated ?2 setae each classical > 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X presence on segments XI to XV not applicable
kingi 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined not applicablenot applicablenot applicablepresence undefined spot/area not applicablepale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. smaller than dark c.s. or approximatively equals not applicable1 pale spot clearly separated or abuted with wing margin 2 pale spots surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableunequal not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segment III presence on segments VIII, IX and X ; presence on segments VII, VIII, IX and X absence on segments XI to XV not applicable
kurensis 1 or more pale spots wholly dark, homogenous ; less than 1/3 pale No pale spots or areas badly defined not applicablenot applicablenot applicableabsence undefined spot/area ; 1 spot well defined absence 1 spot not applicablefused with the r-m crossvein spot or 1 spot absence of at least one of two not applicable1 pale spot clearly separated or abuted with wing margin 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary absence not applicableshort not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals ; unequal not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single ; multiple irregular pits wide opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segment III presence on segments VII, VIII, IX and X presence on segments XI to XIV not applicable
landauae No pale spot cover the r-m crossvein not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableabsence presence 2 functional and 1 rudimentary absence not applicableshort not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 3rd tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV not applicable
langeroni No pale spot cover the r-m crossvein not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicablepyriform absence not applicableapproximatively equals ; unequal not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segments III, IV, V and VI ?presence on segments XI to XIV not applicable
longipennis 1 or more pale spots wholly dark, homogenous Pale spots well defined sizes approximatively equal separated broadly abut wing margin presence 1 spot well defined absence no spot, homogenous cell not applicable2 or 3 spots separated area of pale c.s. smaller than dark c.s. or approximatively equals not applicable1 pale spot clearly separated or abuted with wing margin 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel long not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 3rd tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence presence triangular and moderately swollen ; strongly swollen single wide opening and shallow pit inflated ?1 seta each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X absence on segments XI to XV not applicable
lupicaris 1 or more pale spots more than 1/3 No pale spots or areas badly defined ; Pale spots well defined r3 spot clearly larger than m1 spot separated broadly abut wing margin presence undefined spot/area ; 1 spot well defined absence 1 spot ; pale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. bigger than dark c.s. median margin more narrow than anterior or posterior margin 1 pale spot clearly separated or abuted with wing margin 1 dark spot surrounded by pale area absence absence absence ; almost a wide range of M1 pale absence, no pale spot absence, no pale spot ; almost a wide range of M2 pale absence not applicablelarge shape, elongated spot ; 1 rounded dark spot absence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals ; unequal not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence joined for a short distance (1 or 2 facets) presence both madibles and maxilles absence absence slender or slightly swollen ; triangular and moderately swollen multiple irregular pits not applicableflask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV presence
malevillei No pale spot cover the r-m crossvein not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 3rd tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single narrow opening and deep pit flask shape ?1 seta each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X presence on segments XI to XV not applicable
manchuriensis 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined ; Pale spots well defined sizes approximatively equal ; r3 spot clearly larger than m1 spot separated separated from wing margin absence ; presence no spot, homogenous cell ; undefined spot/area ; 1 spot well defined ?no spot, homogenous cell ; 1 spot ; pale spot over r-m crossvein fused with the m2 spot absence no spot, homogenous cell ; fused with the r-m crossvein spot or 1 spot ?not describedabsence ; 1 pale spot clearly separated or abuted with wing margin ; 2 pale spots fused ; 2 pale spots separated uniform ; 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 1 functional absence not applicableshort oval absence absence ?not applicableabsence not applicablenot applicablenot applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence presence triangular and moderately swollen single wide opening and deep pit flask shape ?2 setae each classical < = 2 ; > 2 presence on segment III ; presence on segments III and V absence on segments VII to X presence on segments XI to XIV ?
marcleti No pale spot cover the r-m crossvein not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence conical short not applicablenot applicablenot applicablenot applicableoval absence not applicableunequal not applicablenot applicableno spines spines on 1st to 3rd tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence presence triangular and moderately swollen single wide opening and shallow pit inflated presence on all short segments 1 seta each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X absence on segments XI to XV not applicable
maritimus 1 or more pale spots wholly dark, homogenous Pale spots well defined sizes approximatively equal separated broadly abut wing margin presence 1 spot well defined absence pale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. smaller than dark c.s. or approximatively equals not applicable2 pale spots fused ; 2 pale spots separated 1 pale spot surrounded by dark area absence absence 1 pale spot in the median part of M1 m1 spot more distal than M1 spot ; M1 and costal spots fused absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel absence not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape presence on all short segments 2 setae each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X presence on segments XI to XV not applicable
maritimus variation 1 or more pale spots wholly dark, homogenous Pale spots well defined sizes approximatively equal separated broadly abut wing margin presence 1 spot well defined absence pale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. smaller than dark c.s. or approximatively equals not applicable2 pale spots fused ; 2 pale spots separated 1 pale spot surrounded by dark area absence absence 1 pale spot in the median part of M1 m1 spot more distal than M1 spot ; M1 and costal spots fused absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel absence not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape presence on all short segments 2 setae each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X presence on segments XI to XIV not applicable
m. paucisensillatus 1 or more pale spots wholly dark, homogenous Pale spots well defined sizes approximatively equal separated broadly abut wing margin presence 1 spot well defined absence pale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. smaller than dark c.s. or approximatively equals not applicable2 pale spots fused ; 2 pale spots separated 1 pale spot surrounded by dark area absence absence 1 pale spot in the median part of M1 m1 spot more distal than M1 spot ; M1 and costal spots fused absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel absence not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape presence on all short segments 2 setae each classical < = 2 ??presence on segments XI to XIV not applicable
m. paucisensillatus variation 1 or more pale spots wholly dark, homogenous Pale spots well defined sizes approximatively equal separated broadly abut wing margin presence 1 spot well defined absence pale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. smaller than dark c.s. or approximatively equals not applicable2 pale spots fused ; 2 pale spots separated 1 pale spot surrounded by dark area absence absence 1 pale spot in the median part of M1 m1 spot more distal than M1 spot ; M1 and costal spots fused absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel absence not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape absence on at least one short segment 2 setae each classical < = 2 ??presence on segments XI to XIV not applicable
minutissimus No pale spot cover the r-m crossvein not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableabsence presence 2 functional and 1 rudimentary absence not applicableshort not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicablespines on 1st and 2nd tarsomere spines on 1st to 3rd tarsomere no spines absence separated widely (more than 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single narrow opening and deep pit ; wide opening and shallow pit inflated ; flask shape ?1 seta each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X absence on segments XI to XV not applicable
montanus 1 or more pale spots more than 1/3 No pale spots or areas badly defined not applicablenot applicablenot applicablepresence undefined spot/area ; 1 spot well defined absence pale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot absence of at least one of two ; area of pale c.s. smaller than dark c.s. or approximatively equals not applicable1 pale spot clearly separated or abuted with wing margin 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence absence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 3rd tarsomere no spines absence joined for a short distance (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single narrow opening and deep pit flask shape ?1 seta each enlarged < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV not applicable
newsteadi 1 or more pale spots more than 1/3 No pale spots or areas badly defined ; Pale spots well defined r3 spot clearly larger than m1 spot separated broadly abut wing margin presence undefined spot/area not applicable1 spot not applicableno spot, homogenous cell ; fused with the r-m crossvein spot or 1 spot area of pale c.s. bigger than dark c.s. median margin wider than anterior or posterior margin 1 pale spot clearly separated or abuted with wing margin 1 dark spot surrounded by pale area absence absence absence ; almost a wide range of M1 pale absence, no pale spot 1 pale spot in median part of M2 ; almost a wide range of M2 pale presence presence large shape, elongated spot absence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals ; unequal not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence joined for a short distance (1 or 2 facets) presence both madibles and maxilles absence absence slender or slightly swollen ; triangular and moderately swollen multiple irregular pits not applicableflask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV ?
nubeculosus 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined not applicablenot applicablenot applicablepresence undefined spot/area not applicablepale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. bigger than dark c.s. anterior, median and posterior margin approximatively equals 2 pale spots fused ; 2 pale spots separated 1 dark spot surrounded by pale area absence presence absence ; almost a wide range of M1 pale absence, no pale spot absence, no pale spot ; almost a wide range of M2 pale absence not applicablenot describedabsence presence 1 functional presence parallel absence elongated curved at third from aperture presence longer than spermatheca not applicableabsence not applicablenot applicablenot applicablenot applicablespines on 1st and 2nd tarsomere spines on 1st to 3rd tarsomere spines on 1st and 2nd tarsomere absence separated widely (more than 2 facets) presence both madibles and maxilles absence absence slender or slightly swollen ; triangular and moderately swollen single ; multiple irregular pits narrow opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segment III presence on segments VIII, IX and X absence on segments XI to XV ?
obsoletus or scoticus 1 or more pale spots more than 1/3 No pale spots or areas badly defined not applicablenot applicablenot applicablepresence undefined spot/area ; 1 spot well defined absence pale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot absence of at least one of two ; area of pale c.s. smaller than dark c.s. or approximatively equals not applicable1 pale spot clearly separated or abuted with wing margin 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence absence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 3rd tarsomere no spines absence joined for a short distance (1 or 2 facets) presence both madibles and maxilles absence absence slender or slightly swollen ; triangular and moderately swollen single narrow opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV not applicable
odiatus 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined not applicablenot applicablenot applicableabsence no spot, homogenous cell not applicableno spot, homogenous cell not applicableno spot, homogenous cell absence of at least one of two not applicableabsence uniform absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary absence not applicableshort not applicablenot applicablenot applicablenot applicableoval absence not applicableunequal not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X presence on segments XI to XIV not applicable
odiatus variation 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined not applicablenot applicablenot applicableabsence no spot, homogenous cell not applicableno spot, homogenous cell not applicableno spot, homogenous cell absence of at least one of two not applicableabsence uniform absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary absence not applicableshort not applicablenot applicablenot applicablenot applicableoval absence not applicableunequal not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segment III ; presence on segments III and V ; presence on segments III and VI ; presence on segments III, V and VI absence on segments VII to X ; presence on segments VII and IX ; presence on segments VII, VIII and IX ; presence on segments VIII, IX and X presence on segments XI to XIV not applicable
pallidicornis 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined not applicablenot applicablenot applicableabsence no spot, homogenous cell not applicableno spot, homogenous cell not applicableno spot, homogenous cell absence of at least one of two not applicableabsence uniform absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 3rd tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen multiple irregular pits not applicableflask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV not applicable
pallidus No pale spot cover the r-m crossvein not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel absence ; short not applicablenot applicablenot applicablenot applicableoval absence not applicable?not applicablenot applicable?spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen ; strongly swollen single narrow opening and deep pit flask shape presence on all short segments 1 seta each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X absence on segments XI to XV not applicable
paolae 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined ; Pale spots well defined sizes approximatively equal ; r3 spot clearly larger than m1 spot separated separated from wing margin presence 1 spot well defined absence 1 spot not applicable2 or 3 spots separated area of pale c.s. smaller than dark c.s. or approximatively equals not applicable2 pale spots separated 1 pale spot surrounded by dark area absence absence 1 pale spot in the median part of M1 m1 spot more distal than M1 spot ; M1 and costal spots separated 1 pale spot in median part of M2 presence presence not applicableabsence presence 2 functional and 1 rudimentary presence parallel absence not applicablenot applicablenot applicablenot applicableoval absence not applicableunequal not applicablenot applicableno spines spines on 1st to 3rd tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence strongly swollen single narrow opening and deep pit flask shape ?2 setae each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X presence on segments XI to XV not applicable
paradisionensis 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined not applicablenot applicablenot applicableabsence no spot, homogenous cell not applicableno spot, homogenous cell not applicableno spot, homogenous cell absence of at least one of two not applicableabsence uniform absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence presence triangular and moderately swollen single wide opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segment III ; presence on segments III and V presence on segments VII and IX presence on segments XIII to XV ; presence on segments XI to XV not applicable
parroti Pale wing, only 1 dark spot not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicablenot applicablepresence 1 functional presence donut absence oval ; elongated kidney shape (heavily curved in the middle) absence ?not applicableabsence not applicablenot applicablenot applicablenot applicablespines on 1st and 2nd tarsomere ; spines on 1st, 2nd and 3rd tarsomere spines on 1st to 3rd tarsomere spines on 1st and 2nd tarsomere ; spines on 1st to 3rd tarsomere absence separated widely (more than 2 facets) presence both madibles and maxilles absence absence slender or slightly swollen multiple irregular pits not applicableflask shape ?1 seta each classical < = 2 presence on segment III presence on segments VIII, IX and X absence on segments XI to XV not applicable
pictipennis 1 or more pale spots more than 1/3 Pale spots well defined sizes approximatively equal separated broadly abut wing margin presence 1 spot well defined absence pale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. smaller than dark c.s. or approximatively equals not applicable1 pale spot clearly separated or abuted with wing margin 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape ?2 setae each classical > 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X presence on segments XI to XV not applicable
picturatus 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined not applicablenot applicablenot applicablepresence no spot, homogenous cell not applicableno spot, homogenous cell not applicablefused with the r-m crossvein spot or 1 spot absence of at least one of two ; area of pale c.s. smaller than dark c.s. or approximatively equals not applicable1 pale spot clearly separated or abuted with wing margin 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel short ; long not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 3rd tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen multiple irregular pits not applicableflask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV not applicable
poperinghensis 1 or more pale spots more than 1/3 No pale spots or areas badly defined not applicablenot applicablenot applicableabsence ; presence no spot, homogenous cell ; undefined spot/area not applicableno spot, homogenous cell ; pale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot absence of at least one of two not applicableabsence ; 1 pale spot clearly separated or abuted with wing margin uniform ; 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence slender or slightly swollen ; triangular and moderately swollen multiple irregular pits not applicableflask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV not applicable
pseudolangeroni No pale spot cover the r-m crossvein not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals ; unequal not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segments III, IV, V and VI ?presence on segments XI to XIV not applicable
pseudopallidus 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined not applicablenot applicablenot applicableabsence no spot, homogenous cell not applicableno spot, homogenous cell not applicableno spot, homogenous cell absence of at least one of two not applicableabsence ; 1 pale spot clearly separated or abuted with wing margin uniform ; 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit inflated ; flask shape ?2 setae each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X presence on segments XI to XV not applicable
pulicaris 1 or more pale spots more than 1/3 No pale spots or areas badly defined ; Pale spots well defined r3 spot clearly larger than m1 spot separated broadly abut wing margin presence undefined spot/area not applicable??no spot, homogenous cell ; fused with the r-m crossvein spot or 1 spot area of pale c.s. bigger than dark c.s. median margin more narrow than anterior or posterior margin 1 pale spot clearly separated or abuted with wing margin 1 dark spot surrounded by pale area absence absence almost a wide range of M1 pale not applicablealmost a wide range of M2 pale absence not applicable1 rounded dark spot absence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals ; unequal not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence joined for a short distance (1 or 2 facets) presence both madibles and maxilles absence absence slender or slightly swollen ; triangular and moderately swollen multiple irregular pits not applicableflask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV absence
punctatus 1 or more pale spots more than 1/3 No pale spots or areas badly defined ; Pale spots well defined r3 spot clearly larger than m1 spot separated broadly abut wing margin presence undefined spot/area not applicable??no spot, homogenous cell ; fused with the r-m crossvein spot or 1 spot area of pale c.s. bigger than dark c.s. median margin wider than anterior or posterior margin 1 pale spot clearly separated or abuted with wing margin 1 dark spot surrounded by pale area absence absence almost a wide range of M1 pale not applicablealmost a wide range of M2 pale presence presence 1 rounded dark spot absence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals ; unequal not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence joined for a short distance (1 or 2 facets) presence both madibles and maxilles absence absence slender or slightly swollen ; triangular and moderately swollen multiple irregular pits not applicableflask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV ?
puncticollis 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined not applicablenot applicablenot applicablepresence undefined spot/area not applicablepale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. bigger than dark c.s. anterior, median and posterior margin approximatively equals 2 pale spots fused ; 2 pale spots separated 1 dark spot surrounded by pale area absence presence absence ; almost a wide range of M1 pale absence, no pale spot absence, no pale spot ; almost a wide range of M2 pale absence not applicablenot describedabsence presence 1 functional presence parallel absence elongated absence ; spermatheca curved at two thirds from aperture absence in length to the spermatheca not applicableabsence not applicablenot applicablenot applicablenot applicablespines on 1st and 2nd tarsomere spines on 1st to 3rd tarsomere spines on 1st and 2nd tarsomere absence separated widely (more than 2 facets) presence both madibles and maxilles absence absence slender or slightly swollen ; triangular and moderately swollen single ; multiple irregular pits narrow opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segment III presence on segments VIII, IX and X absence on segments XI to XV ?
ravus No pale spot cover the r-m crossvein not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence donut long not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals ; unequal not applicablenot applicableno spines spines on 1st to 3rd tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit inflated ?1 seta each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X absence on segments XI to XV not applicable
reconditus No pale spot cover the r-m crossvein not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableabsence presence 2 functional and 1 rudimentary absence not applicableshort not applicablenot applicablenot applicablenot applicableoval presence 2 identical triangular sclerites unequal not applicablenot applicablespines on 1st, 2nd and 3rd tarsomere spines on 1st to 4th tarsomere spines on 1st to 3rd tarsomere ; spines on 2nd and 3rd tarsomere absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence strongly swollen single narrow opening and deep pit flask shape ?2 setae each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X absence on segments XI to XV not applicable
remmi 1 or more pale spots more than 1/3 No pale spots or areas badly defined ; Pale spots well defined r3 spot clearly larger than m1 spot separated broadly abut wing margin presence undefined spot/area not applicablepale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. bigger than dark c.s. anterior, median and posterior margin approximatively equals 1 pale spot clearly separated or abuted with wing margin 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot describedabsence presence 2 functional and 1 rudimentary presence parallel long not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence joined for a short distance (1 or 2 facets) presence both madibles and maxilles absence absence slender or slightly swollen multiple irregular pits not applicableflask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV ?
riebi No pale spot cover the r-m crossvein not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV not applicable
riethi 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined not applicablenot applicablenot applicablepresence undefined spot/area not applicablepale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. bigger than dark c.s. anterior, median and posterior margin approximatively equals 2 pale spots fused ; 2 pale spots separated 1 dark spot surrounded by pale area absence presence absence ; almost a wide range of M1 pale absence, no pale spot absence, no pale spot ; almost a wide range of M2 pale absence not applicablenot describedabsence presence 1 functional presence parallel absence elongated absence ; spermatheca curved at two thirds from aperture absence longer than spermatheca not applicableabsence not applicablenot applicablenot applicablenot applicablespines on 1st and 2nd tarsomere spines on 1st to 3rd tarsomere spines on 1st and 2nd tarsomere absence separated widely (more than 2 facets) presence both madibles and maxilles absence absence slender or slightly swollen ; triangular and moderately swollen single ; multiple irregular pits narrow opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segment III presence on segments VIII, IX and X absence on segments XI to XV ?
riouxi No pale spot cover the r-m crossvein not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableabsence presence 2 functional and 1 rudimentary absence not applicableshort not applicablenot applicablenot applicablenot applicableoval presence 1 triangular and 1 elongated sclerites unequal not applicablenot applicablespines on 1st, 2nd and 3rd tarsomere spines on 1st to 4th tarsomere spines on 1st to 3rd tarsomere ; spines on 2nd and 3rd tarsomere absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence strongly swollen single narrow opening and deep pit flask shape ?2 setae each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X absence on segments XI to XV not applicable
saevus No pale spot cover the r-m crossvein not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableabsence presence 3 functional absence not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicablespherical or oval ?spines on 1st, 2nd and 3rd tarsomere spines on 1st to 4th tarsomere spines on 1st to 3rd tarsomere absence separated widely (more than 2 facets) ; separated narrowly (1 or 2 facets) presence both madibles and maxilles presence absence triangular and moderately swollen single wide opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segments III and V ; presence on segments III and VI ; presence on segments III, V and VI presence on segments VII, VIII, IX and X absence on segments XI to XV not applicable
sahariensis 1 or more pale spots wholly dark, homogenous Pale spots well defined sizes approximatively equal separated broadly abut wing margin presence 1 spot well defined presence pale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. smaller than dark c.s. or approximatively equals not applicable1 pale spot clearly separated or abuted with wing margin 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel long not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 3rd tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence presence triangular and moderately swollen ; strongly swollen single wide opening and shallow pit inflated ?1 seta each classical < = 2 presence on segments III and V presence on segments VII, VIII, IX and X absence on segments XI to XV not applicable
salinarius 1 or more pale spots wholly dark, homogenous Pale spots well defined r3 spot clearly larger than m1 spot separated separated from wing margin presence 1 spot well defined presence 1 spot ; pale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. bigger than dark c.s. anterior, median and posterior margin approximatively equals 2 pale spots fused ; 2 pale spots separated 1 pale spot surrounded by dark area absence absence almost a wide range of M1 pale not applicableabsence, no pale spot ; almost a wide range of M2 pale presence presence not describedabsence presence 1 functional absence not applicableshort oval absence absence ?not applicableabsence not applicablenot applicablenot applicablenot applicableno spines spines on 1st to 3rd tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence presence triangular and moderately swollen single wide opening and deep pit flask shape ?2 setae each classical < = 2 ; > 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X presence on segments XI to XIV ?
santonicus No pale spot cover the r-m crossvein ; 1 or more pale spots wholly dark, homogenous ; less than 1/3 pale No pale spots or areas badly defined not applicablenot applicablenot applicableabsence no spot, homogenous cell not applicableno spot, homogenous cell not applicableno spot, homogenous cell absence of at least one of two not applicableabsence uniform absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicablepresence AND presence of a dark area in the distal part of r5 presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen multiple irregular pits not applicableflask shape ?2 setae each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X presence on segments XI to XV not applicable
santonicus variation No pale spot cover the r-m crossvein ; 1 or more pale spots wholly dark, homogenous ; less than 1/3 pale No pale spots or areas badly defined not applicablenot applicablenot applicableabsence no spot, homogenous cell not applicableno spot, homogenous cell not applicableno spot, homogenous cell absence of at least one of two not applicableabsence uniform absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicablepresence AND presence of a dark area in the distal part of r5 presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen multiple irregular pits not applicableflask shape ?2 setae each classical < = 2 presence on segments III and V presence on segments VII and IX ; presence on segments VII, VIII and IX presence on segments XI to XV not applicable
segnis No pale spot cover the r-m crossvein not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableabsence presence 2 functional and 1 rudimentary absence not applicableshort not applicablenot applicablenot applicablenot applicableoval presence 1 triangular and 1 elongated sclerites unequal not applicablenot applicablespines on 1st, 2nd and 3rd tarsomere spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence strongly swollen single narrow opening and deep pit flask shape ?2 setae each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X presence on segments XI to XIV not applicable
sejfadinei No pale spot cover the r-m crossvein not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableabsence presence 3 functional absence not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableelongated without apical process scale surface spines on 1st to 4th tarsomere spines on 1st to 4th tarsomere spines on 1st to 4th tarsomere absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape ?1 seta each classical < = 2 ?presence on segments VII, VIII, IX and X absence on segments XI to XV not applicable
semimaculatus 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined not applicablenot applicablenot applicableabsence no spot, homogenous cell not applicableno spot, homogenous cell not applicableno spot, homogenous cell ; fused with the r-m crossvein spot or 1 spot absence of at least one of two not applicable1 pale spot clearly separated or abuted with wing margin 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence donut long not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals ; unequal not applicablenot applicableno spines spines on 1st to 3rd tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence strongly swollen single narrow opening and deep pit inflated ?1 seta each classical < = 2 ; > 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X absence on segments XI to XV not applicable
sergenti No pale spot cover the r-m crossvein not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval ; pyriform absence not applicableapproximatively equals ; unequal not applicablenot applicableno spines spines on 1st to 3rd tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X absence on segments XI to XV not applicable
shaklawensis 1 or more pale spots wholly dark, homogenous ; less than 1/3 pale Pale spots well defined r3 spot clearly larger than m1 spot separated broadly abut wing margin presence 1 spot well defined presence pale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. bigger than dark c.s. anterior, median and posterior margin approximatively equals 2 pale spots separated 1 pale spot surrounded by dark area absence absence almost a wide range of M1 pale not applicablealmost a wide range of M2 pale absence not applicablenot describedabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableunequal not applicablenot applicableno spines spines on 1st to 3rd tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape ?2 setae each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV ?
simulator 1 or more pale spots less than 1/3 pale ; more than 1/3 Pale spots well defined r3 spot clearly larger than m1 spot separated broadly abut wing margin presence no spot, homogenous cell not applicablepale spot over r-m crossvein fused with the m2 spot presence fused with the r-m crossvein spot or 1 spot area of pale c.s. smaller than dark c.s. or approximatively equals not applicable1 pale spot clearly separated or abuted with wing margin 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary absence not applicableabsence not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals ; unequal not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape ?2 setae each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X presence on segments XI to XV not applicable
sphagnumensis 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined not applicablenot applicablenot applicableabsence undefined spot/area not applicableno spot, homogenous cell not applicablefused with the r-m crossvein spot or 1 spot absence of at least one of two not applicable1 pale spot clearly separated or abuted with wing margin uniform ; 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 1 functional absence not applicableshort oval absence absence ?not applicableabsence not applicablenot applicablenot applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence strongly swollen single wide opening and shallow pit inflated ?2 setae each classical > 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X presence on segments XI to XV not applicable
stigma Pale wing, only 1 dark spot not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicablenot applicablepresence 1 functional presence parallel absence oval ; elongated finger shape appendage at the bottom absence ?not applicableabsence not applicablenot applicablenot applicablenot applicablespines on 1st and 2nd tarsomere ; spines on 1st, 2nd and 3rd tarsomere spines on 1st to 3rd tarsomere spines on 1st and 2nd tarsomere ; spines on 1st to 3rd tarsomere absence separated widely (more than 2 facets) presence both madibles and maxilles absence absence slender or slightly swollen multiple irregular pits not applicableflask shape ?1 seta each classical < = 2 presence on segment III presence on segments VIII, IX and X absence on segments XI to XV not applicable
subfagineus 1 or more pale spots more than 1/3 Pale spots well defined r3 spot clearly larger than m1 spot separated broadly abut wing margin presence 1 spot well defined absence pale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. bigger than dark c.s. anterior, median and posterior margin approximatively equals 1 pale spot clearly separated or abuted with wing margin 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot describedabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence joined for a short distance (1 or 2 facets) presence both madibles and maxilles presence absence slender or slightly swollen multiple irregular pits not applicableflask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV ?
subfasciipennis 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined not applicablenot applicablenot applicableabsence no spot, homogenous cell not applicableno spot, homogenous cell not applicableno spot, homogenous cell absence of at least one of two not applicableabsence ; 1 pale spot clearly separated or abuted with wing margin 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 3rd tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen multiple irregular pits not applicableflask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV not applicable
subfasciipennis Marocco Variety 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined not applicablenot applicablenot applicableabsence no spot, homogenous cell not applicableno spot, homogenous cell not applicableno spot, homogenous cell absence of at least one of two not applicableabsence ; 1 pale spot clearly separated or abuted with wing margin 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 3rd tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV not applicable
submaritimus 1 or more pale spots wholly dark, homogenous Pale spots well defined r3 spot clearly larger than m1 spot separated broadly abut wing margin presence 1 spot well defined absence pale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. smaller than dark c.s. or approximatively equals not applicable2 pale spots fused ; 2 pale spots separated 1 pale spot surrounded by dark area absence absence 1 pale spot in the median part of M1 m1 spot more distal than M1 spot ; M1 and costal spots fused absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel absence not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape presence on all short segments 2 setae each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X presence on segments XI to XIV not applicable
tauricus No pale spot cover the r-m crossvein not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableabsence presence 3 functional absence not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableelongated without apical process smooth surface spines on 1st to 4th tarsomere spines on 1st to 4th tarsomere spines on 1st to 4th tarsomere absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape ?1 seta each classical < = 2 ?presence on segments VII, VIII, IX and X absence on segments XI to XV not applicable
tbilisicus No pale spot cover the r-m crossvein not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableabsence presence 2 functional and 1 rudimentary absence not applicableshort not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen multiple irregular pits not applicableflask shape ?1 seta each classical < = 2 presence on segment III absence on segments VII to X presence on segments XI to XV not applicable
truncorum 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined not applicablenot applicablenot applicablepresence no spot, homogenous cell not applicableno spot, homogenous cell not applicableno spot, homogenous cell absence of at least one of two ; area of pale c.s. smaller than dark c.s. or approximatively equals not applicable1 pale spot clearly separated or abuted with wing margin 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel absence not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence strongly swollen single wide opening and shallow pit flask shape ?2 setae each classical < = 2 presence on segments III and V presence on segments VII and IX presence on segments XI to XV not applicable
univittatus 1 or more pale spots wholly dark, homogenous ; less than 1/3 pale Pale spots well defined sizes approximatively equal separated separated from wing margin presence 1 spot well defined absence pale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot area of pale c.s. smaller than dark c.s. or approximatively equals not applicable1 pale spot clearly separated or abuted with wing margin 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel short not applicablenot applicablenot applicablenot applicableoval absence not applicableunequal not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape ?2 setae each classical < = 2 presence on segments III, IV, V and VI presence on segments VII, VIII, IX and X presence on segments XI to XV not applicable
vexans No pale spot cover the r-m crossvein not applicablenot applicablenot applicablenot applicablenot applicablenot applicableno spot, homogenous cell not applicableno spot, homogenous cell not applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicablenot applicableabsence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel short ; long not applicablenot applicablenot applicablenot applicableoval absence not applicableapproximatively equals not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen multiple irregular pits not applicableflask shape ?2 setae each classical < = 2 presence on segments III and V presence on segments VII and IX presence on segments XI to XV not applicable
vidourlensis 1 or more pale spots wholly dark, homogenous No pale spots or areas badly defined not applicablenot applicablenot applicableabsence ; presence no spot, homogenous cell ; undefined spot/area not applicablepale spot over r-m crossvein fused with the m2 spot absence fused with the r-m crossvein spot or 1 spot absence of at least one of two ; area of pale c.s. smaller than dark c.s. or approximatively equals not applicable1 pale spot clearly separated or abuted with wing margin 1 pale spot surrounded by dark area absence absence absence absence, no pale spot absence, no pale spot absence not applicablenot applicableabsence presence 2 functional and 1 rudimentary presence parallel absence ; short not applicablenot applicablenot applicablenot applicableoval absence not applicableunequal not applicablenot applicableno spines spines on 1st to 4th tarsomere no spines absence separated narrowly (1 or 2 facets) presence both madibles and maxilles absence absence triangular and moderately swollen single wide opening and shallow pit flask shape ?1 seta each classical < = 2 presence on segments III and V presence on segments VII and IX presence on segments XI to XV not applicable